"Medusa," a deadly virus becomes a worldwide epidemic. In order to escape from this deadly virus, a handful of people are chosen to be put into a cold sleep, laying in a capsule hoping for the future cure. Kasumi, a teenage girl is one of the 160 chosen for this procedure, and is guided to a Cold Sleep Capsule Center (CSCC) inside an ancient castle. Understanding that it is hardly possible, Kasumi goes to sleep still anticipating for a reunion with her twin sister Shizuku, who also is infected with the virus. As Kasumi and the others awake, they notice that the CSCC is not as they remembered. Just like the story of "Sleeping Beauty," the castle is covered with thorn, and the awaken are attacked by unknown creatures and monsters! How long were they asleep? Where did the monsters come from? What has happened to the world? Abandoned in the midst of an enigma, the escapade of the seven survivors begins...

The Fairytale World is in trouble. Its princesses and their respective worlds are disappearing, causing a ripple effect in the human world where their stories are popular. In order to save the Fairytale World, the Queen sends three magic animals, Sei, Dai, and Ryoku, to the human world with magic gems to find three girls who can become the "Super Miracle Idols," the princesses Snow White, Cinderella, and Kaguya-hime. Those "princesses" end up being three little girls: Yukimori Ringo, Takashiro Layla, and Sasahara Natsuki. But the gems transform them into older singing superstars, and after their accidental debut at the singer Wish's concert, they become known as "Little Princesses," or "LilPri." Now they must collect Happiness Tones from humans in order to restore the Fairytale World.

The film's story is set in the Doll Kingdom where dolls reside. Happiness Charge PreCure are asked to help by a women named Tsumugi when the sudden appearance of Saiarks begin wrecking havoc upon the Kingdom. However all is not what it seems with the mysterious man Black Fang dwelling in the shadows.

After years of fruitless war between the four realms of Gamindustri (Planeptune, Lastation, Lowee and Leanbox) over Share energy, the source of their strength based on how much their people have faith in their goddesses, the four CPUs that rule over them have finally signed a friendship treaty. The treaty bans any attempt at claiming Share energy through military force, in hopes of bringing peace and prosperity to their worlds. Yet, a month after the treaty, Neptune, the CPU Goddess of Planeptune, spends her time goofing off and playing games rather than doing her job, leaving her land's Shares plummeting. Choujigen Game Neptune The Animation follows Neptune and her friends' attempts at raising Shares, while dealing with an external threat that could spell the end of both the Goddesses and Gamindustri itself...