At a CD store, Ichika Hoshino hears a song by Miku that she has never heard before. On the monitor, she sees a "Hatsune Miku" in a form she has never seen before, and couldn't help say it out loud, "Miku!?" Miku was surprised at the voice, her eyes meet Ichika's, but she disappears shortly after. Later, after a live performance on the street, the Miku that Ichika saw before appears on her smartphone. Miku looks sad and downcast, and when Ichika quietly asks Miku about her story, she tells Ichika that there are people to whom she wants to deliver her songs, but no matter how much she sings, her songs are not reaching them. Miku, who saw Ichika singing to the hearts of many people at her live performances, thought that if she learns more about Ichika, she can do the same, and came to Ichika. Ichika smiles at Miku and says, "If it's all right with me." And a new story of Hatsune Miku and the boys and girls begins—.

Second installment of the movie.

Movie sequel to .

Hoping to unite their countries with an alliance, The Empress of Austria arranges her daughter, the delightful yet spoiled Marie Antoinette, to marry the crown prince of France, Louis XVI. Upon arriving in Versailles, the Austrian princess meets Oscar Francois de Jarjayes, Captain of the Royal Guards—a seemingly dashing young man—but to Antoinette's astonishment, he is in fact a woman! The youngest daughter of a noble family with a prominent lack of a male heir, Oscar was raised as a boy to continue the family's military legacy. Suffocated by the rigid rules associated with her new position and longing for companionship, Antoinette immediately takes a liking to this intriguing young woman, and wastes no time befriending her. However, as the childish new queen foolishly abuses her power, Oscar is conflicted between maintaining her loyalty to the royal family and addressing the growing concern of poverty among the common people. Recalling the events of the French Revolution, Versailles no Bara depicts the struggles of two women unjustly burdened by family expectations. Yet, despite being forced to deny their identities at the expense of personal happiness, they march on to become key players in the annals of history.