Adaptation of the Hashira Training Arc. The Hashira, the Demon Slayer Corps' highest ranking swordsmen and members. The Hashira Training has begun in order to face the forthcoming battle against Muzan Kibutsuji. Each with their own thoughts and hopes held in their hearts, a new story for Tanjiro and the Hashira begins.

The introduction of 7G Network technology promised miraculous technological advancements; however, the aftermath of its launch plunged Japan into chaos. From drastic changes to the terrain to bizarre mutations among people, the way of life is overwritten with new rules. The town of Agano is one of the few remaining settlements, and its citizens aged 21 and above have transformed into animals. Shizuru Chikura, a high school student from Agano, is desperately searching for her friend, Youka Nakatomi. Coming across a picture in a newspaper placing Youka in Ikebukuro, Shizuru starts packing for a trip. But with public transportation effectively shut down, her options are limited. Fortunately, Tairo Zenjirou, a senile old conductor and the only human adult in town, comes to the rescue when he teaches her how to operate an abandoned train. With her destination set, Shizuru ventures off on a journey to find her missing friend. On the day of departure, however, her other classmates—Nadeshiko Hoshi, Reimi Kuga, and Akira Shinonome—decide to tag along at the last minute as they embark on this one-track adventure that might expose them to dangers far surpassing anything they have encountered in Agano.

Sousuke Kaburaya, an impoverished detective, meets Sara, a princess from another world with magical powers. They start living together, and Sara quickly adjusts her life to modern Japan.

The story is set in an alternate world called Yamauchi, ruled by the Yatagarasu clan, who are able to transform between human and crow forms. Yukiya, a beautiful and eccentric Yatagarasu boy, is chosen to serve the young prince. While encountering various events in the conspiracy-filled imperial court, he forms a strange master-servant relationship with the young prince.